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In prokaryotes, such as E. coli, in which no nuclear envelope exists, translation begins before transcription is completed. Figure 11.21 shows a length of an E. coli chromosome. An RNA polymerase is visible on the DNA, transcribing a gene. The messenger RNA, still being synthesized, can be seen extending away from the DNA.Attached to the messenger RNA are about a dozen ribosomes. Since translation starts at the end of the messenger that is synthesized first (5 ), an initiation complex can form and translation can begin shortly after transcription begins. As translation proceeds along the messenger, its 5 end will again become exposed, and a new initiation complex can form.The occurrence of several ribosomes translating the same messenger is referred to as a polyribosome, or simply a polysome ( g. 11.22). In prokaryotes, most messenger RNAs contain the information for several genes. These RNAs are said to be polycistronic ( g. 11.23). (Cistron, another term for gene, is de ned in chapter 12.) Each gene on the messenger RNA is translated independently: each has a ShineDalgarno sequence for ribosome recognition (see g.11.12) and an initiation codon (AUG) for fMet.The ribosome that completes the translation of the rst gene may or may not continue to the second gene after dissociation.The translation of any gene follows all the steps we have outlined.

that is, that the voltage across an element is directly proportional to the current ow through it R is the value of the resistance in units of ohms ( ), where 1 = 1 V/A (214)

The resistance of a material depends on a property called resistivity, denoted by the symbol ; the inverse of resistivity is called conductivity and is denoted by the symbol For a cylindrical resistance element (shown in Figure 220), the resistance is proportional to the length of the sample, l, and inversely proportional to its cross-sectional area, A, and conductivity, v= l i A

diamond tool cutting material and on the right a cross-sectional view of the tool and the workpiece [41]

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Peptide bond formation on the ribosome between N-formyl methionine and phenylalanine. The bond attaching the carboxyl end of the rst amino acid to its tRNA is transferred to the amino end of the second amino acid. The rst tRNA is now uncharged, whereas the second tRNA has a dipeptide attached.

(215)

tides that mimic anticodons to recognize the stop codons: proline-alanine-threonine in RF1 and serineproline-phenylalanine in RF2. In this molecular mimicry, a protein mimics the shape of a nucleic acid in order to function properly. The next base in the messenger RNA past the stop codon is usually an adenine, required for ef cient termination.After the release factors act, with the hydrolysis of a GTP, the ribosome has completed its task of translating mRNA into a polypeptide. Final release of all factors and dissociation of the two subunits of the ribosome take

Physical resistors with resistance R Typical materials are carbon, metal film Circuit symbol i-v characteristic

It is often convenient to de ne the conductance of a circuit element as the inverse of its resistance The symbol used to denote the conductance of an element is G, where G= 1 siemens (S) R where 1 S = 1 A/V (216)

The McGraw Hill Companies, 2001

Thus, Ohm s law can be restated in terms of conductance as: I = GV (217)

An initial model was developed based on indentation fracture mechanic analysis According to Scattergood, fracture initiation plays a central role for ductile-regime machining A critical penetration depth, dc, for fracture initiation was derived by Blackley and Scattergood [41] as follows:

2

(a) EF-G s translocation of the ribosome converts it from a pretranslocational state (P and A sites occupied) to a posttranslocational state (E and P sites occupied). The uncharged transfer RNA in the E site is then ejected. (b) A seethrough model of the 70S ribosome of E. coli with transfer RNAs in the A, P, and E sites. The structure was determined by cryoEM mapping, an electron microscopic technique using rapidly frozen specimens. The position of the messenger RNA is shown, as well as the stalk of the 50S subunit (St) and one of the polypeptides of the large subunit, L1. ([b] Courtesy of

Exponential i-v characteristic (semiconductor diode)

Joachim Frank, Howard Hughes Medical Institute.)

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