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Chain termination at the ribosome. One of two release factors recognizes a nonsense codon in the A site. In this case, RF1 recognizes UAG. The complex then falls apart, releasing the peptide.

+ vS(t) + _ + Vmax RL vL(t) vS(t) + _

Figure 860 Equivalent circuit for the one-sided limiter (diode on)

(a) Protein synthesis at a polysome. Nascent proteins exit from a tunnel in the 50S subunit. Messenger RNA is being translated by the ribosomes while the DNA is being transcribed. (b) A messenger RNA from the midge, Chironomus tentans, showing attached ribosomes and nascent polypeptides emerging from the ribosomes. Note the 5 end of the messenger RNA at the upper right (small peptides). Magni cation 165,000 . ([b] Courtesy of S. L. McKnight and

(33)

Figure 861 Equivalent circuit for the one-sided limiter (doide off)

If, on the other hand, the source voltage is such that RL vS (t) < Vmax rS + R L then D1 is an open circuit and the load voltage is simply vL (t) = RL vS (t) rS + R L (827) (826)

O. L. Miller, Jr.)

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The equivalent circuit for this case is depicted in Figure 861 The analysis for the negative branch of the circuit of Figure 858 can be conducted by analogy with the preceding derivation, resulting in the waveform for the two-sided clipper shown in Figure 862 Note how the load voltage is drastically clipped by the limiter in the waveform of Figure 862 In reality, such hard clipping does not occur, because the actual diode characteristic does not have the sharp on-off breakpoint the ideal diode model implies One can develop a reasonable representation of the operation of a physical diode limiter by using the piecewise linear model 2 Piecewise linear diode model To avoid unnecessary complexity in the analysis, assume that Vmax is much greater than the diode offset voltage, and therefore assume that V 0 We do, however, consider the nite diode resistance rD The circuit of Figure 859 still applies, and thus the determination of the diode

A GGAGGU AUG UGA AGGAGGU AUG UGA AGGAGGU AUG UGA 5 Shine-Dalgarno sequences Figure 11.23 3

8

tan b = ( f/R) cos q From equation (33) and the aforementioned triangle in Figure 341, lets X = f/R sin b = cos b =

10 5 Volts 0 5 10 0 02 04 t (s) 06 08 1

A prokaryotic polycistronic mRNA. Note the several Shine-Dalgarno sequences for ribosomal attachment and the initiation and termination codons marking each gene.

on-off state is still based on whether [RL /(rS + RL )]vS (t) is greater or less than Vmax When D1 is open, the load voltage is still given by vL (t) = RL vS (t) rS + R L (828)

vL(t)

Figure 11.21 A polysome (i.e., multiple ribosomes on the same strand of mRNA). Each ribosome is approximately 250 units across. Also visible in this illustration are DNA and RNA polymerase. (Reproduced courtesy of Dr. Barbara Hamkalo, International

Figure 863 Circuit model for the diode clipper (piecewise linear diode model)

(34) (35)

Review of Cytology, (1972) 33:7, g. 5. Copyright by Academic Press, Inc., Orlando, Florida.)

When D1 is conducting, however, the corresponding circuit is as shown in Figure 863 The primary effect the diode resistance has on the load waveform is that some of the source voltage will reach the load even when the diode is conducting This is most easily veri ed by applying superposition; it can be readily shown that the load voltage is now composed of two parts, one due to the voltage Vmax , and one proportional to vS (t): vL (t) = RL rS rD RL Vmax + vS (t) rD + (RL rS ) rS + (rD RL ) (829)

It may easily be veri ed that as rD 0, the expression for vL (t) is the same as for the ideal diode case The effect of the diode resistance on the limiter circuit is depicted in Figure 864 Note how the clipping has a softer, more rounded appearance

mRNA 5 Nascent polypeptide (early)

5 10 0 01 02 03 04 05 06 t (s) 07 08 09

Figure 864 Voltages for the diode clipper (piecewise linear diode model)

Nascent polypeptide (late) (b)

Fig 341:

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